Early Pleistocene ground beetles (Coleoptera: Carabidae) from the Ookui Formation in Nagano Prefecture, central Japan, and their biogeographical and paleoenvironmental significance

M. Hayashi

1998

Article

The Quaternary Research 37:117-129

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Taxa discussed in this work:

• Agabus optatus
• Chlaenius gebleri
• Hemicarabus

Taxonomy:

• Agabus optatus belongs to Agabus - Sharp (1884)
• Chlaenius gebleri belongs to Chlaenius - Ganglbauer (1891)
• Hemicarabus belongs to Carabidae - Hayashi (1998)
• Apotomopterus belongs to Carabidae - Hayashi (1998)